Description
The mandrill’s facial colouration — the most vivid colouration of any mammal, the ridged blue of the maxillary swellings and the red of the nose and lips intensifying with testosterone levels — is a reliable indicator of male dominance status. The dominant male of a large mandrill group in Lopé National Park carries the most saturated facial pigmentation of any male in the troop. This image was made during a field study of the Lopé mandrill population conducted by the Max Planck Institute, the dominant male habituated to researcher presence through years of observation. The close-range portrait was made at 200mm from a distance of 6 metres, the focal length and distance creating the perspective compression that emphasises the facial colouration against the softly blurred forest background. The yellow beard — fully developed in dominant males — frames the blue-and-red facial display. The image required months of habituation work by the field team.
